University of Texas MD Anderson Cancer Center

Distinguished Research Professor Emeritus Dalhousie Univ

School of Biological Sciences, University of Manchester

Professor of Genetics, Friedrich Schiller University Jena

Professor of Botany, University of British Columbia

Dept. of Cell Biology & Anatomy, University of Calgary

Professor of Biological Sciences, University of Alberta

University of Padova

Class of 1955 Endowed Professor of Biology, Indiana University, Bloomington

Professor in Biology and Director of the Scottish Oceans Institute, University of St Andrews