Department of Zoology, University of Cambridge

Professor of Biology, University of St Andrews

Canada Excellence Research Chair, University of Alberta, UNSW

University of Liverpool

Professor of Behavioural Ecology, University of Bristol

The University of Sydney

Edward Grey Institute, Department of Biology, University of Oxford

Professor of Animal Welfare, University of Bern, Switzerland

Department of Veterinary Medicine, University of Cambridge

Emeritus Professor of Mathematics and Biology, University of Bristol